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Mac Hippenhammer also took first-team reps with the wideouts.
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Trace McSorley signals to the sideline.
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Lamont Wade worked at safety during the defensive series.
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Penn State's first-team offensive line saw Chasz Wright, Mike Menet, Connor McGovern, Steven Gonzalez and Ryan Bates, right to left.
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Wideout Brandon Polk.
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Nick Scott intercepted a last-second heave downfield.
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Defensive coordinator Brent Pry celebrated the play.
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Head coach James Franklin addressed the team after practice.
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Amani Oruwariye spoke to reporters following practice.
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James Franklin answered questions after practice. The Nittany Lions have two more spring practices, including Saturday's Blue-White Game.